Nina, a wyndian princess, and her guardian Cray are searching everywhere for Nina's missing elder sister, Elina. While traveling to her last known whereabouts they suffer an accident. On her way to a nearby town, Nina meets Ryu, a seemingly amnesiac young man. This meeting triggers a series of events that will involve them with the godly race known as The Endless, immortal dragons with the power to shape the world itself.

Closely follows the storyline of the original PlayStation game.